Simulated effects of increasing atmospheric CO, and changing climate on the successional characteristics of Alpine forest ecosystems

Possible effects of changing climate and increasing CO, on forest stand development were simulated using a forest succession model of the JABOWA/FORET type. The model was previously tested for its ability to generate plausible community patterns for Alpine forest sites ranging from 220 m to 2000 m a.s.l., and from xeric to mesic soil moisture conditions. Each model run covers a period of 1000 yrs and is based on the averaged successional characteristics of 50 forest plots with an individual size of 1/12 ha. These small forest patches serve as basic units to model establishment, growth, and death of individual trees. The simulated CO, scenario assumes linear climate change as atmospheric CO, concentration increases from 310 p1/1 to 620 pl/l and finally to 1340 pl/l. Direct effects of increasing CO, on tree growth were modeled using treering and growth chamber data. The simulation experiment proved to be a useful tool for evaluating possible vegetation changes that might occur under C0,-induced warming. On xeric sites from the colline to the high montane belt, the simulated climate change causes drastic soil water losses due to elevated evapotranspiration rates. This translates into a significant biomass decrease and even to a loss of forest on xeric low-elevation sites. Biomass gains can be reported from mesic to intermediate sites between 600 and 2000 m a.s.1. Increasing CO, and warming alters the species composition of the simulated communities considerably. In today's montane and subalpine belt an invasion of deciduous tree species can be expected. They outcompete most conifers which in turn may migrate to today's alpine belt. Some of these changes occur as early as 40 yrs after climate begins to change. This corresponds to a mean annual warming of 1.5"C compared with today's mean temperatures.